As a side note, Cataglyphis ants primarily use stride integration, but they fall back Excellence is not a skill it is an attitude shirt on visual cues. In their native environment, there are very few visual cues. Other ants have native environments with many visual cues and depend primarily on visual cues but fall back on stride integration. Yet other ants are entirely visual and do not appear to do any stride integration. So the particular navigational strategy varies with the ant and the environment that selected for her abilities And then there are the tandem runners, like Temnothorax, some Camponotus, some Pachycondyla, and even more. The details can vary in the tandem running use across these ant taxa to a new location without the use of a trail. The leader walks a short distance and waits to be touched by the follower.
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Meanwhile, the follower Excellence is not a skill it is an attitude shirt encodes the visual landscape as she sweeps her head from side to side. After learning the entire route, she can return on her own and even take novel paths showing the destination as opposed to the entire route was actually communicated, and she memorized that location. There are variants of tandem running, like in Pachycondyla where the follower learns the path while being carried without any chemicals being used along the trail. In general, these tandem behaviors show the use of both ant memory as well as visual capability.